Scottish Rock Garden Club Forum

General Subjects => Alpines => Topic started by: TheOnionMan on October 17, 2010, 02:24:06 AM

Title: Aquilegia saximontana
Post by: TheOnionMan on October 17, 2010, 02:24:06 AM
Searching around on the forum, there have been questions about the valid identity of Aquilegia saximontana.  I couldn't find a photo on the current SRGC Forum that I could say was A. saximontana without question, although did find one photo on the old archived SRGC forum by Graham Nicholls that show the real species.
http://www.srgc.org.uk/discus/messages/283/30636.html#POST15434  (scroll to find it, or use the next link below)
http://www.srgc.org.uk/discus/messages/283/30941.jpg

Recently I came across some good resources that should help illustrate what this Colorado endemic actually looks like, simply as a forum resource.

US Forest Service page on 5 blue columbines, starting with A. saximontana.  Click each photo for an enlarged view.
http://www.fs.fed.us/wildflowers/beauty/columbines/aquilegia_saximontana.shtml

Uploaded are several photos showing two different plants of A. saximontana growing in Panayoti Kelaidis' troughs.  The closeup view shows the hooked spurs (uncommon for American blue species, but common enough for columbine species elsewhere), the narrowish cup with diffuse color shift from blue to white or cream, and delicately flared petals at full anthesis.  Notice too, the small acute leaf dentations.  The flowers are tiny. 

Also uploaded is a fine line drawing (scanned) of this species, and A. laramiensis (endemic to Wyoming).  The photos are from Gentes Herbarum, Aquilegia - The Cultivated and Wild Columbines, by Philip A. Munz, March 1946, Vol. VII, 150 pp.

Flora of North America
http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=233500115
Title: Re: Aquilegia saximontana
Post by: Diane Clement on October 17, 2010, 09:34:22 AM
Some interesting links, thanks Mark.  The seed exchanges often receive and list "Aquilegia jonesii x saximontana" but I remember Bob Nold saying that he didn't think this hybrid exists.  So are these reputed crosses just A saximontana or a different hybrid?  Has anyone grown them on?
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on October 17, 2010, 01:35:42 PM
Some interesting links, thanks Mark.  The seed exchanges often receive and list "Aquilegia jonesii x saximontana" but I remember Bob Nold saying that he didn't think this hybrid exists.  So are these reputed crosses just A saximontana or a different hybrid?  Has anyone grown them on?

Diane, regarding the putative Aquilegia jonesii x saximontana hybrid, I have yet to see a plant or image convincing me that the hybrid exists.  Given the well known promiscuity of the genus, I imagine that garden seed collected from either species, when other columbines species/hybrids are present, will likely result in hybrid plants not true to the species.  It is more likely that anything going around as "Aquilegia jonesii x saximontana" is a mongrel plant with all sorts of parentage possibilities.  The name has been distributed in seed exchanges for so many years that surely by now the name is meaningless, a mixed bag of columbine genetics, with any of the dozens of dwarf Aquilegias so popular among rock gardeners contributing to the mélange.
Title: Re: Aquilegia saximontana
Post by: Lesley Cox on October 17, 2010, 09:36:59 PM
Some years ago I grew seed under this hybrid name. The resultant plants looked more like A. flabellata than anything else, and not the 'Nana' form either, but quite large and vigorous.
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on October 17, 2010, 11:43:35 PM
Some years ago I grew seed under this hybrid name. The resultant plants looked more like A. flabellata than anything else, and not the 'Nana' form either, but quite large and vigorous.

Why does this not surprise me!  ;)
Title: Re: Aquilegia saximontana
Post by: Great Moravian on October 18, 2010, 12:28:22 PM
The following links are for comparison.
Aquilegia glandulosa 'Jucunda'  
http://www.picturetrail.com/sfx/album/main/153947/199252071
Extremely short spurs. The plant as all plants usually cultivated denoted so in reality belongs to Aquilegia flabellata without any connection to Aquilegia glandulosa. The latter can be found at http://www-sbras.nsc.ru/win/elbib/atlas/flora/408.html and http://www.botanicgardensblog.com/wp-content/uploads/2009/07/aquilegia-glandulosa-smallest2.jpg and http://www.amaranthus.ru and http://pisum.bionet.nsc.ru/kosterin/plants/ranunculaceae/glandulosa.htm
Aquilegia pyrenaica subsp. discolor
http://www.picturetrail.com/sfx/album/main/153947/199815274
White spurs noticeable.
Aquilegia saximontana
http://www.picturetrail.com/sfx/album/main/153947/199482438
Title: Re: Aquilegia saximontana
Post by: Maggi Young on October 18, 2010, 12:53:49 PM
I understood that Aquilegia saximontana,native to the Rocky Mountains of Colorado, was  a diminutive plant, growing little more than 10 cms ( 4 inches) high in the wild, though perhaps a little more in cultivation (as is quite commonly found to be the case with mountain flowers)
The flowers are blue with white and the spurs distinctly hooked at the tips and the sexual parts discretely hidden.

I believe that the plant is as shown in the July 2010 #7 of International Rock Gardener
- the tiny stature, foliage colour and short, curling spurs  convince me!

These links are to various  USA sites ....
The curling spurs are clearly illustrated here :
http://www.efloras.org/object_page.aspx?object_id=40998&flora_id=1
 this is the description....
http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=233500115


http://www.fs.fed.us/wildflowers/beauty/columbines/aquilegia_saximontana.shtml

http://calphotos.berkeley.edu/cgi/img_query?enlarge=0000+0000+1002+0444

Title: Re: Aquilegia saximontana
Post by: TheOnionMan on October 18, 2010, 04:24:11 PM

I believe that the plant is as shown in the July 2010 #7 of International Rock Gardener http://www.srgc.org.uk/logs/logdir/2010Jul291280437029IRG7_July2010.pdf
- the tiny stature, foliage colour and short, curling spurs  convince me!

These links are to various  USA sites ....
The curling spurs are clearly i llustreated here :

Regarding the photo in IRC #7, sorry but I'm not convinced, it does not look like A. saximontana to me, but rather like a dwarf form of A. flabellata (nana, var. pumila, other names and combinations) or a hybrid of it.  Aquilegia flabellata var. pumila, like many European and Asian aquilegia, also have tightly curled spurs.  In my opinion, the plant in that photo has flowers of the wrong "character", they look plump or rotund, incurved, of heavier texture and striated, deeper blue-purple coloration, and the cream lamina strongly demarcated from the blue portion, versus a diffuse color transition.  It doesn't look right to me.

Some further links for comparison:

Aquilegia flabellata (nana, pumila, yezoense, Mini-Star, pumila 'Mini-Star')
http://1003gardens.blogspot.com/2009/04/aquilegia-flabellata-nana-yezoense-easy.html

http://2.bp.blogspot.com/_TAp_ZAaoiUM/SeUSYFymSMI/AAAAAAAACH4/JKJo8pK7XqE/s1600-h/Aquilegia+flabellata+nana-Yezoense.jpg

http://www.stauder.net/bildearkiv/Aquilegia%20flabellata%20nana%202%20%206,3.jpg

http://www.pflanzen-vielfalt.de/popup_image.php?pID=910842

http://commons.wikimedia.org/wiki/File:Aquilegia_flabellata_var_pumila1.jpg

NARGS page on trough, A. flabellata 'Nana' part way down:
http://www.nargs.org/nargswiki/tiki-index.php?page=troughs

as A. akitensis, a synonym of flabellata
http://www.zahradnictvolimbach.sk/skalnicky/aquilegia_akitensis.jpg

var. pumila
http://www.wildgingerfarm.com/images/Aquilegiaflabellata.jpg

var. pumila (in N. Japanese Alps)
http://www.flickr.com/photos/akira_o/918992319/in/set-72157600833042281/
http://www2.yamanashi-ken.ac.jp/~yohnishi/personal/mountains/photos/plants/salpsn/odamaki.kita.jpg
Title: Re: Aquilegia saximontana
Post by: Maggi Young on October 18, 2010, 07:59:32 PM
Interesting links.... none of which look like the plant in IRG 7  :-X

 [attach=1]
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on October 18, 2010, 09:18:20 PM
Interesting links.... none of which look like the plant in IRG 7  :-X


That's the problem, nothing quite looks like the plant in IRG 7, particularly saximontana. ::)
Title: Re: Aquilegia saximontana
Post by: Maggi Young on October 18, 2010, 09:34:58 PM
This illustration from efloras.org, Flora of North America shows a flower form that is like the IRG 7 pic....
[attach=1]

 With photographes there is always the problem of comparing plants at differing stages of growth, in or out of character, with varying colour casts in the camera etc.

I will say that the description of foliage,flower etc I have seen matches the description of wild plants and given a provenance of Colorado seed rather than European or Japanese, I am more inclined to find it to be saximontana than any other. 
Title: Re: Aquilegia saximontana
Post by: tonyg on October 18, 2010, 11:13:34 PM
Interesting links.... none of which look like the plant in IRG 7  :-X

  (Attachment Link)
I'm with Mark - it looks like A flabellata (nana) as I have grown it ... and as Google sees it!
Pretty though ... as are the pics of Panayotis plants, seed please!
Title: Re: Aquilegia saximontana
Post by: Maggi Young on October 18, 2010, 11:25:47 PM

I see the flower profile of saximontana as being much rounder, as shown in the illustration from efloras, also, I  rather think the plant shown originated from Colorado seed  ::)

I'll agree to disagree.
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on October 19, 2010, 04:16:23 AM
The Flora of North America line drawing of A. saximontana shows an important detail; acute sepals.  However, like all line drawings, they capture one moment in time, on a single individual plant.  One must observe full anthesis on the flowers to see what the flower disposition is, and on A. saximontana the sepals are widely spreading, giving a light airy look to the flowers.

Yes Maggi, we can agree to disagree. Regarding "given a provenance of Colorado seed rather than European or Japanese", it is important to note that the photo in IRG 7 depicts a garden grown plant, and any of number of factors could be at play; seed mix-up, seed contaminated with other species when cleaned or as provided by the seller, plant labeling mixup, digital photo labeling "after the fact" misidentification, etc.

Meanwhile to help Aquilegia fans understand key characteristics of true Aquilegia saximontana, I prepared a composite image showing 4 flowers of A. saximontana, 3 taken from the US Forest Service page showing 3 different plants, and the 4th from Panayoti Kelaidis' plant of known provenance. Key things to note: the sepals are acute (rather than widely rounded as in A.f. pumila or nana), and at full anthesis, the sepals spread widely.  The lamina ends (or "cup") is narrowish, not rotund as in A. f. pumila/nana.  The spurs on A. saximontana splay outwards then bend inwards at the tip... the hook at the end is mild (a very distinctive and characteristic look), whereas in A. f. pumila/nana the spurs angle inwards (converging) and strongly coiled at the tips to almost 360 degrees.  To help illustrate the spur differences, I put together two detail drawings from the aforementioned Gentes Herbarum publication, showing the differences in spurs; slight bend in saximontana, strong coil bend in flabellata, the strong coil bend visible in the IRC 7 image.  And not specifically described, but true in every image I've seen to date, on A. saximontana the blue color blends in diffuse manner to the cream or white cup, whereas in A. f. pumila/nana the color change is a sharp demarcation as in the IRC 7 image.

So, I remain firmly convinced that the plant shown in IRC 7 shows a flabellata form or hybrid, and looks nothing like A. saximontana.  I'm dedicating some time on this, as this species (saximontana) is nearly legendary in terms of misidentifications for decades.  I well remember seeing true Aquilegia saximontana in Colorado in the 1980s for the first time and thinking, "wow, that's what it really looks like", then realizing that nothing under this name in the seedexes (at that time) was even close to being true.  As I find more photos showing plants that look like true A. saximontana I shall post them here as reference.

By the way Tony, you have a good eye, as does google ;-)
Title: Re: Aquilegia saximontana
Post by: Great Moravian on October 19, 2010, 12:42:07 PM
Quote
This illustration from efloras.org, Flora of North America shows a flower form that is like the IRG 7 pic...
You are right. The image is obviously misleading. It is a general problem of botanical illustration which exaggerates certain distinctive characters but forgets others not important for a given purpose.
In Aquilegia saximontana the spurs are usually divergent at flower bases whereas in the cultivars of Aquilegia flabellata the spurs are usually parallel or convergent at flower bases. It is the consequence of petal shape. The petal limb is deeply bulgy in Aquilegia flabellata, but not in Aquilegia saximontana and Aquilegia glandulosa which of course is a large-flowered species.
Title: Re: Aquilegia saximontana
Post by: Great Moravian on October 19, 2010, 01:59:21 PM
Interesting links.... none of which look like the plant in IRG 7
The Aquilegia flabellata var. pumila at the Japanese
http://ptech.cocolog-nifty.com/blog/2008/04/post_869f.html
It certainly looks like the plant in IRG 7.
Title: Re: Aquilegia saximontana
Post by: maggiepie on October 19, 2010, 03:03:40 PM
I am finding this thread very interesting.

Here are a couple of pics of seedlings of mine.

Aquilegia saximontana seedling



Aquilegia jonesii x saximontana

Lastly, an A. flabellata bought from a nursery

I don't know what the first three actually are but I like them a lot.
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on October 19, 2010, 03:45:32 PM
Interesting links.... none of which look like the plant in IRG 7
The Aquilegia flabellata var. pumila at the Japanese
http://ptech.cocolog-nifty.com/blog/2008/04/post_869f.html
It certainly looks like the plant in IRG 7.

Agreed.  Last night I too was looking at this Japanese blog site, lots of good pics of Aquilegia.  In the following link, way down near the bottom is a photo of what the grower purports to be the true A. saximontana.  Translating the page with Google Translate (which does a poor job), what I can make out is that this grower tried for 10 years to get the true plant, but in the photo he shows in 2006, believes he has the true plant.  I agree it looks right.  To facility finding the photo easier, in fair use I have uploaded a screen capture of that image (see 1st photo).  Here again we see the distinctly divergent spurs, mild hook to the spurs, and acute sepals.
http://sainohana.cocolog-nifty.com/blog/2006/05/index.html

Found a second page by the same Japanese grower in 2010, can't be sure from the translation, but I believe these pictures taken 4 years later might be seedling plants, and thus subject to hybridization.  Again, in fair use, I put two screen captures of the flowers side by side for comparison, showing characteristic spurs.
http://ptech.cocolog-nifty.com/blog/2010/04/post-3eb0.html

As an aside, here's another link on this same blog to A. flabellata var. pumila:
http://ptech.cocolog-nifty.com/blog/2009/04/aquilegia-flabe.html

And, just a curiosity, but a couple double-flowered dwarf Aquilegia that were found on Rebun Island, Japan :o
http://sainohana.cocolog-nifty.com/blog/2006/05/post_ad14.html
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on October 19, 2010, 04:03:30 PM
I am finding this thread very interesting.
Here are a couple of pics of seedlings of mine.

Aquilegia saximontana seedling

Aquilegia jonesii x saximontana

Lastly, an A. flabellata bought from a nursery

I don't know what the first three actually are but I like them a lot.

Helen, the first two look like A. flabellata (dwarf forms) or dwarf flabellata hybrid to me, not saximontana, although I agree that they're really cute.  The last one is an A. flabellata dwarf form as you've identified.  Hard to know about the one identified as jonesii x saximontana... really a choice little thing.  The degree of pubescence on the flowers and spurs seems stronger than jonesii alone, and the strength of the hooked spurs, indicate other species influences in my opinion.
Title: Re: Aquilegia saximontana
Post by: maggiepie on October 19, 2010, 06:23:56 PM
Mark, I'm not at all sure that the first two are flabellata. I have lots of those and they all grow into a lovely little mound, these ones are only about half the height of the flabellata and the leaves look different. If they flower at the same time next year I will have a really good squizz at them.

I wish I had tried to get seed from them.
I don't suppose you know a way of getting rid of wee harlequin stinkbugs ?

Title: Re: Aquilegia saximontana
Post by: Maggi Young on October 19, 2010, 06:49:44 PM
Has no-one else noticed the distinct differences in the "saximontana" plants - form flowers and foliage- between the first two of McMark's photos of Kelaidis plants and the third?
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on October 19, 2010, 08:02:42 PM
Mark, I'm not at all sure that the first two are flabellata. I have lots of those and they all grow into a lovely little mound, these ones are only about half the height of the flabellata and the leaves look different. If they flower at the same time next year I will have a really good squizz at them.


Helen, the first two look flabellata-esque to me, including the foliage, that is to say they are probably dwarf flabellata hybrids as suggested.  There could be saximontana genes in there too, just that I strongly doubt pure saximontana. 

Many forms of flabellata exist under a variety of names (nana, var. pumila, yezoense, akitensis, akitensis kurilensis, flabellata v. kurilensis), and as obtained via the seed exchanges from open-pollinated garden sources, well, we all know what happens to Aquilegia when grown from OP garden seed, you get aquilegia stew.  When I was a younger fellow, I was focused on columbines, and grew a large number of them, and had many versions of plants I would attribute to "flabellata lineage", and it was near impossible to get some species true from such OP seed.  I also grew a number of the dwarf columbines from collected sources, such as bertolonii and discolor, all which were charming, but in the open garden, with many columbines tending to be short-lived and seeding around, after a few years I could no longer be certain of their purity, they started stewing.

Title: Re: Aquilegia saximontana
Post by: mark smyth on October 20, 2010, 12:16:20 PM
That Japanese person has some very nice ?miniature doubles
Title: Re: Aquilegia saximontana
Post by: Great Moravian on October 20, 2010, 02:06:00 PM
Plants are variable in leaves. Compare the few-leaved A. flabellata var. pumila found in Rebun at
http://www.rebun-island.jp/en/hana/index.html
---
(http://www.srgc.org.uk/smf/index.php?action=dlattach;topic=6126.0;attach=248780;image)
Spurs parallel, A. flabellata
---
(http://www.srgc.org.uk/smf/index.php?action=dlattach;topic=6126.0;attach=248782;image)
Spurs parallel-convergent, A. flabellata
---
(http://www.srgc.org.uk/smf/index.php?action=dlattach;topic=6126.0;attach=248784;image)
Leaves completely different, the plant is usually cultivated as A. bertolonii, which is not correct too, or A. pyrenaica, which is possible.
---
(http://www.srgc.org.uk/smf/index.php?action=dlattach;topic=6126.0;attach=248786;image)
Spurs parallel, A. flabellata
---
(http://www.srgc.org.uk/smf/index.php?action=dlattach;topic=6126.0;attach=248790;image)
Spurs divergent, A. saximontana
---
(http://www.srgc.org.uk/smf/index.php?action=dlattach;topic=6126.0;attach=248792;image)
Spurs divergent, A. saximontana
Title: Re: Aquilegia saximontana
Post by: Maggi Young on October 20, 2010, 02:11:51 PM
I refer again to McMark's scan of the drawings.... and to the two plants  shown..... the last looks like a saximontans but the first two pictures are surely, by his reasoning, laramiensis?  ::)

Great Moravian...... I can see what you mean to say.... but the photos you show are displaying a different meaning to the one I understand of paralllel, for a start....... parallel means being the same distant apart along the length......  :-X
Title: Re: Aquilegia saximontana
Post by: Great Moravian on October 20, 2010, 02:25:28 PM
I refer again to McMark's scan of the drawings.... and to the two plants  shown..... the last looks like a saximontans but the first two pictures are surely, by his reasoning, laramiensis?  ::)
I am not certain which image should depict A. laramiensis. The flowers if it are usually whitish.
Quote
Great Moravian...... I can see what you mean to say.... but the photos you show are displaying a different meaning to the one I understand of paralllel, for a start....... parallel means being the same distant apart along the length......  :-X
Despite I am a mathematician, I don't expect plants to be geometric objects. All terms in botany are merely approximate.
Title: Re: Aquilegia saximontana
Post by: Maggi Young on October 20, 2010, 02:30:58 PM
Quote
Despite I am a mathematician, I don't expect plants to be geometric objects. All terms in botany are merely approximate.

 Exactly!!
Title: Re: Aquilegia saximontana
Post by: Maggi Young on October 20, 2010, 02:33:24 PM
Quote
I am not certain which image should depict A. laramiensis.
I rather thought the image labelled laramiensis??
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on October 20, 2010, 02:48:58 PM
I refer again to McMark's scan of the drawings.... and to the two plants  shown..... the last looks like a saximontans but the first two pictures are surely, by his reasoning, laramiensis?  ::)


Don't be fooled by the line drawing and apparent size difference, once again, the scan depicts just 1 drawing of 1 plant each, it cannot take into account variability.  Little separates saximontana and laramiensis, mostly separated on flower color and geographical location, with other minor differences.  Both species can be 5-25 cm tall in the wild.  There are some very nice photos on the web of this species, maybe I'll post links, no photos of laramiensis have been posted on this topic yet.

From FONA key
Sepals and spurs white or nearly so; Wyoming.      Aquilegia laramiensis
Sepals and spurs blue; Colorado.                          Aquilegia saximontana

I call your Roll Eyes and raise by another Roll Eyes  ::) ::)
Title: Re: Aquilegia saximontana
Post by: Maggi Young on October 20, 2010, 03:04:19 PM
McMark.... I'm not talking about scale.... look at the drawings, look at the leaf differences... now look at the pictures of the first plant in PK's trough... the leaves have their central section well clear, of the others, making a very distinct elongated triangular outline for the whole.....


I would be more than astonished if these different aquilegia were NOT as variable as 99.9 percent of other plants..... it is only humans who seek absolute delineations of species, to satisfy our assorted desires for listing and boxing things..... it would be nice to see that some cognisance was being taken of natural variability and that the various problems, mistakes, hiccups etc etc that have been listed earlier can apply just as well to one photo as another.
Life is rarely clear cut, botany is certainly no different...... there is little of exactitude in such matters. 

 


Title: Re: Aquilegia saximontana
Post by: TheOnionMan on October 20, 2010, 03:23:27 PM
Moving on, I will gather up some nice photos I found of A. laramiensis and post those links here, a species that could be legitimately confused with saximontana, except for flower color and more subtle differences.
Title: Re: Aquilegia saximontana
Post by: Maggi Young on October 20, 2010, 03:40:08 PM
Aquilegia laramiensis is very little grown  in UK gardens, as far as I can tell.... I'm sure many would like to have some links to pictures of this delicate ivory beauty, McMark


 I'll give you a start....http://www.kadel.cz/flora/Images/WebSize/220.jpg

http://www.fs.fed.us/r2/projects/scp/assessments/aquilegialaramiensis.pdf
Title: Re: Aquilegia saximontana
Post by: Great Moravian on October 20, 2010, 03:41:59 PM
In A. laramiensis the flowers are hanging, small, whitish, and spurs are approximately parallel.
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on October 21, 2010, 04:18:51 AM
Maggi, you grabbed the first two A. laramiensis links I had on my list.  Still gathering, but getting late here and I'm tired. 

So for tonight, with fair use I scanned and posted a photo I had forgotten about, Aquilegia saximontana in Rare Plants of Colorado, Second Edition, 1997, The Colorado Native Plant Society.  It's a good characteristic image, showing the divergent spurs.

Also with fair use, I scanned and posted a drawing of Aquilegia saximontana from Land Above the Trees, A Guide to American Alpine Tundra, by Anne Zwinger and Beatrice Willard, 1972.  I still consider this one of the very best books on alpine plant habitats, the pages filled with charming pencil drawings that capture the essence of the subject plant, often showing the flowers and plant in various stages of articulation.  Here again we see acute sepals, narrowish cup, and divergent spurs.

The 3rd and last fair use scan is Aquilegia saximontana from Rocky Mountain Alpines, part of Alpines '86, Second Interim International Rock Garden Conference, June 28 to July 2, 1986; Boulder, Colorado.  Sorry, the scan is a bit pixelated, but the image was rather small, but you can still see the unique shape of the flower and spurs.
Title: Re: Aquilegia saximontana
Post by: Great Moravian on October 21, 2010, 10:09:44 AM
http://www.fs.fed.us/wildflowers/beauty/columbines/aquilegia_laramiensis.shtml
http://davesgarden.com/guides/pf/showimage/152108/
http://calphotos.berkeley.edu/cgi/img_query?stat=BROWSE&query_src=photos_flora_sci&where-genre=Plant&where-taxon=Aquilegia+laramiensis&title_tag=Aquilegia+laramiensis
Title: Re: Aquilegia saximontana
Post by: Great Moravian on October 21, 2010, 11:03:37 AM
In my opinion, the identity of A. laramiensis and A. saximontana was sufficiently clarified.
The former is not a showy species, the latter is prone to hybridization. I suggest to
discuss the identity of two showy and non-hybridizing species, namely
http://www.srgc.org.uk/smf/index.php?action=dlattach;topic=6126.0;attach=248784;image
and
http://www.picturetrail.com/sfx/album/main/153947/199815274
The two are in my cultivation since 1975.
The former is distributed as A. bertolonii, A. pyrenaica or A. jonesii x saximontana,
the latter usually as A. discolor.
Surprisingly, the two don't cross despite of their belonging to one species if the former is
A. pyrenaica subsp. pyrenaica and the latter A. pyrenaica subsp. discolor.
Title: Re: Aquilegia saximontana
Post by: Graeme Strachan on October 21, 2010, 12:53:42 PM
Does anyone know what I really have? I posted this in april 2009.
http://www.srgc.org.uk/smf/index.php?topic=3329.msg89842#msg89842 (http://www.srgc.org.uk/smf/index.php?topic=3329.msg89842#msg89842)

         regards

            Graeme Strachan
Title: Re: Aquilegia saximontana
Post by: Great Moravian on October 21, 2010, 01:13:41 PM
Does anyone know whay I really have? I posted this in april 2009.
http://www.srgc.org.uk/smf/index.php?topic=3329.msg89842#msg89842 (http://www.srgc.org.uk/smf/index.php?topic=3329.msg89842#msg89842)
A Japanese cultivar of Aquilegia flabellata var. pumila in my guess.
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on October 21, 2010, 01:15:44 PM
Does anyone know whay I really have? I posted this in april 2009.
http://www.srgc.org.uk/smf/index.php?topic=3329.msg89842#msg89842 (http://www.srgc.org.uk/smf/index.php?topic=3329.msg89842#msg89842)
A Japanese cultivar of Aquilegia flabellata var. pumila in my guess.

Moravian, I just tried posting and it said another message was already posted... great minds think alike ;D

Graeme, I second the opinion above, it looks like a cute little A. flabellata var. pumila (or hybrid of it).  Looking around the web, I have found many lovely desirable forms of A. flabellata var. pumila, here's a particularly attractive one.
http://www.aquabiom.sk/images/katalog/1196715239_1_big.jpg?rand=33856806
Title: Re: Aquilegia saximontana
Post by: Great Moravian on October 21, 2010, 03:47:38 PM
In reality, we are disgusting people who are happy apparent growers of presumed botanical rarities. I wonder whether anybody got happier after determination of her-his treasure. Replacing of rarities using either hybrids or other easily cultivable plants is a common procedure making living easier.
Title: Re: Aquilegia saximontana
Post by: Diane Clement on October 21, 2010, 04:10:08 PM
In my opinion, the identity of A. laramiensis and A. saximontana was sufficiently clarified.
The former is not a showy species, the latter is prone to hybridization.

But is it "prone to hybridisation"?  Has anyone ever grown or even seen the fabled A jonesii x saximontana (and it wasn't A flabellata or similar).  I'm not sure it exists.
Title: Re: Aquilegia saximontana
Post by: Graeme Strachan on October 21, 2010, 09:32:46 PM
Moravian/Onion Man - Thank you for identifying my plant. I am happy that more knowledgeable people have put me right rather than living in ignorance and expounding the problem when I give away its offspring.  8)

        regards

             Graeme Strachan
Title: Re: Aquilegia saximontana
Post by: maggiepie on October 21, 2010, 10:04:25 PM


But is it "prone to hybridisation"?  Has anyone ever grown or even seen the fabled A jonesii x saximontana (and it wasn't A flabellata or similar).  I'm not sure it exists.

Am wondering if anyone would recognize it if it does exist. ???
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on October 21, 2010, 11:34:47 PM
Moravian/Onion Man - Thank you for identifying my plant. I am happy that more knowledgeable people have put me right rather than living in ignorance and expounding the problem when I give away its offspring.  8)

        regards

             Graeme Strachan

Excellent Graeme, my inspiration for this thread is to provide enough information and verifiable images on the universally misidentified Aquilegia saximontana, to help lessen perpetual redistribution of plants under the wrong name.  Moravian has been most helpful in this effort too, and as well, has shed some light on other often-misidentified species too, that species like A. bertolonii, pyrenaica, discolor, are also mired in confusion, an unfortunate state of Aquilegia for decades. But the focus here is on A. saximontana, it's closest ally A. laramiensis, and the species that most often usurps the good name of saximontana; namely A. flabellata and its forms.

I see that you had no reply to your earlier posting on your Aquilegia ID.  Just after your post, there was another post with an image of Aquilegia akitensis (synonym of A. flabellata).  One thing we haven't discussed here yet, is some of the foliar differences between saximontana and flabellata, but I believe A. flabellata helps reveal itself by its foliar characteristics as well.
Title: Re: Aquilegia saximontana
Post by: Olga Bondareva on October 27, 2010, 11:01:55 AM
I am disappointed by this topic. I understand many of my aquilegias are under wrong names.

This one came like A. saximontana
(http://cs4145.vkontakte.ru/u6450879/93407084/x_86b9ccb8.jpg)

This one like A. bertolonii
(http://cs4145.vkontakte.ru/u6450879/93407084/x_40d04252.jpg)

This - A. discolor
(http://cs4289.vkontakte.ru/u6450879/92164489/x_988744fa.jpg)

A. flabellata v. pumilio
(http://cs723.vkontakte.ru/u6450879/20107304/x_b06097ed.jpg)

And A. flabellata v. pumilio Rosea
(http://cs723.vkontakte.ru/u6450879/20107304/x_469968c0.jpg)

But now I can not say they are truly recognized.  ???
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on October 27, 2010, 04:21:43 PM
I am disappointed by this topic. I understand many of my aquilegias are under wrong names.

But now I can not say they are truly recognized.  ???

Olga, don't be too disappointed, you have some beautiful columbines there.  The fact that Aquilegia are promiscuous and garden grown seed cannot be relied upon for species purity goes way back... I attach a scan taken from Gentes Herbarum, 1946, with some discussion on Aquilegia flabellata... it is rather telling regarding the confusion that already ensued back in the early 20th century.

Your first white and blue one (not saximontana) looks similar to the link that Moravian posted above, here it is again:
http://www.picturetrail.com/sfx/album/main/153947/199815274
...a plant that goes around as A. discolor, A. pyrenaica subsp. discolor.

Your last one is an exquisite rose form, very nice.  I notice the name "pumilio" is used sometimes, there is a var. pumila... can't tell how large your pink flabellata is, but it is a gem.  Your plants nicely photographed as usual.
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on October 28, 2010, 01:09:20 PM
Just came across this photo of Aquilegia saximontana in a past NARGS Rock Garden Quarterly, Volume 66 Number 3 Summer 2008, the photo courtesy of Denver Botanic Garden.  Here again we see the light airy foliage and flowers with divergent spurs. 
Title: Re: Aquilegia saximontana
Post by: Gunilla on October 28, 2010, 02:09:24 PM
Very interesting topic.  Like you, Olga I have a lot of misnamed Aquilegias and this tiny blue and white one looks similar to your first.
Wrongly named A. saximontana.
Title: Re: Aquilegia saximontana
Post by: Great Moravian on October 29, 2010, 01:04:09 PM
I am disappointed by this topic. I understand many of my aquilegias are under wrong names.

This one came like A. saximontana
(http://cs4145.vkontakte.ru/u6450879/93407084/x_86b9ccb8.jpg)
A. pyrenaica subsp. dicolor
***
This one like A. bertolonii
(http://cs4145.vkontakte.ru/u6450879/93407084/x_40d04252.jpg)
A. pyrenaica subsp. pyrenaica  or A. bertolonii. The latter is usually a coarser plant.
***
This - A. discolor
(http://cs4289.vkontakte.ru/u6450879/92164489/x_988744fa.jpg)
Cultivar of A. flabellata var. pumila
***
A. flabellata v. pumilio
(http://cs723.vkontakte.ru/u6450879/20107304/x_b06097ed.jpg)
Cultivar of A. flabellata var.pumila
***
And A. flabellata v. pumilio Rosea
(http://cs723.vkontakte.ru/u6450879/20107304/x_469968c0.jpg)
Cultivar of A. flabellata var pumila

But now I can not say they are truly recognized.  ???
Title: Re: Aquilegia saximontana
Post by: cohan on October 31, 2010, 02:11:22 AM
an interesting topic..
how far do aquilegias go with their promiscuity? if for example  i had a couple of large garden cultivars, and one of the north american native oranges, (i do) and were to get wild collected seed of one of the western miniatures, such as jonesii or others, would these be sufficiently different to be safe, or will they still cross among themselves?
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on October 31, 2010, 02:53:24 AM
an interesting topic..
how far do aquilegias go with their promiscuity? if for example  i had a couple of large garden cultivars, and one of the north american native oranges, (i do) and were to get wild collected seed of one of the western miniatures, such as jonesii or others, would these be sufficiently different to be safe, or will they still cross among themselves?

Unless one were to keep plants isolated and covered to prevent insect pollination, then the flowers selfed, it is my belief that they will start hybridizing right away.  I have found that all of the North American red and red-orange types will hybridize readily with other species, to such a degree that one needs to be growing plants from wild seed often to keep them going in the garden.  You'll get close look-alikes from seed, but one loses confidence in only a few years of growing seedlings after seeing first hand the hybrid shift.

I've never been fortunate enough to grow A. jonesii, and it could be that some highly specialized and genetically isolated species are not so quick to produce interspecific hybrids, but in my experience open-pollinated Aquilegia seedlings will very quickly turn into columbine soup, the soup can be muddled and of diminished quality, or be rather good one with pretty plants.  On the NARGS Forum, I posted a photo essay of a garden visit of a friend's garden in Central, Massachusetts, there are some photos of Aquilegia shown there, such as A. barnebyii (exquisite, grown from wild collected seed), but I make mention of the owner's older part of the garden in which Aquilegia have seeded about for years and blended into surprisingly pretty mix of colors, all obviously hybrids.
http://nargs.org/smf/index.php?topic=373.0

What I did not show in that essay, are the many plants of A. "canadensis" in his garden... while they look like canadensis at quick first glance, when you look closely one realizes these are hybrids, intergrading with dominant vulgaris types.  In many gardens it seems that A. flabellata is strong genetically, an overriding presence in subsequent generations of seedlings, this has been my experience through the years.  Moravian indicated that A. pyrenaica ssp. pyrenaica and A. pyrenaica ssp. discolor don't cross (amongst themselves), but I suspect they still cross with other species, Aquilegia being the most promiscuous of promiscuous genera.
Title: Re: Aquilegia saximontana
Post by: Olga Bondareva on October 31, 2010, 02:38:55 PM
Thank you Mark and Great Moravian!

So there are no A. saximontana between mine. Could you (or somebody) show me A. scopulorum? I have 2 years old plants with very blue leaves. But they still have not flowered.

And may I ask you about some more Aquilegias?
This one I get from seed taken at Tajikistan.

(http://cs10003.vkontakte.ru/u6450879/31156622/x_eec5edb7.jpg)  (http://cs10003.vkontakte.ru/u6450879/31156622/x_c5b3c5e7.jpg)
It is a favorit of slugs and fungi.  :(

This one is named like A. laramiensis.

(http://cs9312.vkontakte.ru/u6450879/20107304/x_e3780590.jpg)

It is small and leaves are purplish.
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on October 31, 2010, 02:48:53 PM
The American Aquilegia scopulorum is less confused than saximontana, as scopulorum is a small bun of foliage with variably colored flowers with exceptionally long spurs, the flowers tending to face upwards.  Here are some good links:

Aquilegia scopulorum
http://www.rmrp.com/Photo%20Pages/AA/Aquilegia%20scopulorum%20100DPI.htm

http://www.fs.fed.us/wildflowers/beauty/columbines/aquilegia_scopulorum.shtml

Nice views here, scroll down after gallery loads:
http://aplantaday.blogspot.com/2010_07_01_archive.html

http://plants.usda.gov/java/profile?symbol=AQSC
http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=233500116
Title: Re: Aquilegia saximontana
Post by: Olga Bondareva on October 31, 2010, 02:55:36 PM
Thanks Mark!

Leaves looks similar. My plants didn't flowered so I didn't saw spurs.  :)
Title: Re: Aquilegia saximontana
Post by: Maggi Young on October 31, 2010, 04:15:32 PM
There will be a lovely A. scopulorum ssp. perplexans picture by Joyce Carruthers in the International Rock Gardener for November.
Title: Re: Aquilegia saximontana
Post by: cohan on October 31, 2010, 05:32:13 PM
an interesting topic..
how far do aquilegias go with their promiscuity? if for example  i had a couple of large garden cultivars, and one of the north american native oranges, (i do) and were to get wild collected seed of one of the western miniatures, such as jonesii or others, would these be sufficiently different to be safe, or will they still cross among themselves?

Unless one were to keep plants isolated and covered to prevent insect pollination, then the flowers selfed, it is my belief that they will start hybridizing right away.  I have found that all of the North American red and red-orange types will hybridize readily with other species, to such a degree that one needs to be growing plants from wild seed often to keep them going in the garden.  You'll get close look-alikes from seed, but one loses confidence in only a few years of growing seedlings after seeing first hand the hybrid shift.

good to know, mark..i haven't got any of the little blues, yet, but would like to at some point..
there is a small patch of some large garden hybrid slowly seeding around in a dry area under an old spruce--the original plant has 'black' flowers, but i saw a white flowering seedling this year, first to flower, though the original has  been there many years; the orange flowered plants are originally from b.c. (around 30 years ago, but overgrown by junipers)and are maybe 50metres or more from the others, i just noticed one orange flowered seedling 10m or so from the parent..

when i go for a blue i will go for wild seed, and maybe deadhead...
Title: Re: Aquilegia saximontana
Post by: Graham Catlow on October 31, 2010, 07:40:22 PM
I have been watching this from the start and it just gets more complicated.
The reason for my interest is that I bought A. scopulorum in the Spring but it is not scopulorum. In fact it looks nothing like scopulorum. It flowered but I didn't photograph it. It looks closer to saximontana but I will have to wait until next year for a photo.
I also bought A. pyrenaica and A. canadensis both of which may not be what I thought they were.
Will just have to wait until the Spring.
Graham
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on November 01, 2010, 04:17:00 AM
There will be a lovely A. scopulorum ssp. perplexans picture by Joyce Carruthers in the International Rock Gardener for November.

I'm a bit confused, I saw a nice photo of A. scopulorum var. scopulorum in the International Rock Gardener for October, is the November issue already out?

Just a point of nomenclature, the name A. scopulorum "ssp. perplexans" Clokey 1938 is a synonym for A. scopulorum.  The "ssp. perplexans" was named for a race in Southern Nevada (not Utah) with more color variability than was thought normal, but soon put into synonymy with the type species.  There does exist A. scopulorum var. calcarea, which is recognized by USDA but not Flora of North America, for a race with petioles that are glandular-pubescent.  There is also a recent Aquilegia scopulorum var. goodrichii S.L.Welsh 2003, which I know almost nothing about, except that it is on the endangered list.
http://plants.usda.gov/java/profile?symbol=AQSC
http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=233500116

Title: Re: Aquilegia saximontana
Post by: Maggi Young on November 01, 2010, 09:43:04 AM
The November issue of IRG wil not be online until 26th November. 
The photo that will be included is of that multi coloured race you mention.
The  ssp. perplexans is kept by the author and I am happy to go with that.
The information as to the name changes /synonomy are easily found by those seeking further information but the relevance is to a population of variously coloured plants.   
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on November 01, 2010, 12:17:11 PM
Thanks Maggi, I look forward to seeing the next IRG issue with that photo, along with more buckwheats.

I'm out for the day, but tonight will post what is written about A. scopulorum ssp. perplexans in the Gentes Herbarum monograph on Aquilegia, it explains the color variation and I think readers will be interested.
Title: Re: Aquilegia saximontana
Post by: maggiepie on November 01, 2010, 12:24:05 PM
Well, this thread has excited my lust for aquilegias again.
Am going to sow all the old seeds I have that I didn't get around to, some might still be viable.
I have several packets of 'saximontana' seed from several seed exchanges.
 ;D ;D ;D
Title: Re: Aquilegia saximontana
Post by: Lesley Cox on November 01, 2010, 08:35:42 PM
I haven't seen mention of the Italian (?) A. bertolonii in this thread. I have no picture unfortunately but in my experience, it does seem always to come true from seed, even though I have had many others growing nearby. It seems to be the American species that are most prone to hybridizing.
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on November 01, 2010, 11:11:58 PM
I haven't seen mention of the Italian (?) A. bertolonii in this thread. I have no picture unfortunately but in my experience, it does seem always to come true from seed, even though I have had many others growing nearby. It seems to be the American species that are most prone to hybridizing.

Lesley, Great Moravian mentioned that what is often distributed A. bertolonii is typically incorrect, akin to A. pyrenaica
http://www.srgc.org.uk/smf/index.php?topic=6126.msg169677#msg169677
http://www.srgc.org.uk/smf/index.php?topic=6126.msg169554#msg169554

I haven't focused on species such as these, because when I started this topic I wanted to specifically highlight A. saximontana as among the most misidentified columbine around, and also give context with associated species and likely misnomers (thus the focus on the ubiquitous flabellata).  Moravian has provided broader context to Aquilegia identifications and misidentification, which has been most helpful, particularly in light of the nearly mythical creature A. jonesii x saximontana.
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on November 02, 2010, 02:17:26 AM
Thanks Maggi, I look forward to seeing the next IRG issue with that photo, along with more buckwheats.

I'm out for the day, but tonight will post what is written about A. scopulorum ssp. perplexans in the Gentes Herbarum monograph on Aquilegia, it explains the color variation and I think readers will be interested.

As mentioned, here is the description of Aquilegia scopulorum ssp. perplexans (Clokey 1938) from the Gentes Herbarum 1946 monograph Aquilegia The Cultivated and Wild Columbines by Philip A. Munz.  The subspecies is included under the species A. scopulorum as a synonym.  The description is:  The subspecies perplexans of Charleston Peak, southern Nevada, seems to have no differentiating character except some red in the flowers of some of the plants.  Utah plants range from blue, blue-purple to white for sepals and bluish to yellowish petals.  Since the Charleston Mountain plants have these colors, or add, in some cases, red to give red, blue, blue and white, red and white, red and yellow, white, yellow, they seem too indefinite to stand as a separate subspecies".

I would love to see some of those reported color variants, particularly a red and white form, as pictures I've seen, and plants I've grown, are generally in the blue to blue and white range of things. I've seen pics of nice pink flowered forms.

It is curious that "ssp. perplexans" is wholly absent, even as a synonym, from the USDA North American flora plant profiles, from ITIS (Integrated Taxonomic Information System), and from Flora of North America, the latter only recognizing A. scopulorum. My floras of both Nevada and Utah do not cite any name "perplexans".  I've seen such examples a few times, where a named plant species, variety or subspecies, early in its trajectory is deemed invalid, subsequently getting left out of commonly cited synonymy possibly because of it's speedy nullification.  However, such names often live a long and prosperous perpetuated life in horticulture regardless of long standing taxonomic definitions to the contrary.
Title: Re: Aquilegia saximontana
Post by: Great Moravian on November 02, 2010, 10:01:15 AM
This one I get from seed taken at Tajikistan.
(http://cs10003.vkontakte.ru/u6450879/31156622/x_eec5edb7.jpg)  (http://cs10003.vkontakte.ru/u6450879/31156622/x_c5b3c5e7.jpg)
Aquilegia lactiflora - vicaria - pseudovicaria - tianschanica -karatavica - darwazii alliance discussed in a parallel thread
http://www.srgc.org.uk/smf/index.php?topic=6155.0
Probably Aquilegia vicaria because spurs are longer to sepals.
Title: Re: Aquilegia saximontana
Post by: Great Moravian on November 02, 2010, 11:41:18 AM
The problem of distinguishing between Aquilegia bertolonii and Aquilegia pyrenaica subsp. pyrenaica is difficult.
There are no differences in the descriptions and keys in Flora europaea and Flore de France. The species overlap in all character values.
For example, spurs are usually hooked and sometimes curved or straight in the former but usually curved or straight and sometimes hooked in the latter.
Bertoloni himself denoted specimens of Aquilegia bertolonii as Aquilegia pyrenaica.
Schott described Aquilegia bertolonii and compared Aquilegia pyrenaica as follows.
...
Aquilegia Bertolonii S. Foliis infimis infra pilosulis, supra  
glabratis
, lobulis linearibus subacutatis, divergentibus; caule  
gracili, ramuloso, inferne petiolisque pilosulo l. glabrato, apicem
versus dense viscide-puberulo; floribus magnis; sepalis oblongo-
lanceolatis, acuminatis, ciliolatis, postice sparsepilosulis, antice
glabratis ; petalorum lamina oblonga, sepalis multo breviore,
apice subtruncato-rotundata, extus puberula, intus excepta  
summitate glabra, ciliolata; calcare uncinato, puberulo, cras-
siusculo, lamina sua subaequilongo ; genitalibus petalis paulo
brevioribus; cyamiis . . . . . . . .  

...
Aquilegia pyrenaica DC. foliis infimis infra pilosulis, supra  
glabratis, lobulis oblongo-obovatis, truncato-rotundatis, retusis,
subdistantibus; caule gracili, inferne petiolisque pilosulo, apicem
versus viscido-puberulo; floribus magnis; sepalis oblongo-
ovatis, subcuspidatis, ciliolatis, extus pilosulis, intus glabris;
petalorum lamina oblonga, sepalis multo breviore, apice ro-
tundata, utrinque praecipue apicem versus dense-puberula,
ciliolata, calcare rectiusculo, puberulo, valde attenuato, la-
mina sua multo longiore; genitalibus petalis multo brevioribus;  
cyamiis viscide-puberulis rostro subaequilongo auctis.

...
The differences in orange are not mentioned by later botanists.
In reality, the sepals are not oblong-ovate in the latter but really substantial in comparison to the former.
Compare the tapering sepals of Aquilegia bertolonii
http://luirig.altervista.org/cpm/albums/bot-006/aquilegia-bertolonii11936.jpg
and the substantial sepals in magnificent forms of Aquilegia pyrenaica subsp. pyrenaica
http://www.fotonatura.org/galerias/fotos/usr22713/11999063cm.jpg
but not as substantial in other forms
http://4.bp.blogspot.com/_dhmGoQyou-c/S0tzCpbIu_I/AAAAAAAABQM/GM91LIyr77Y/s400/Aquilegia+pyrenaica+blog.jpg
So the differences in brown are of statistical nature.
Spurs in the two specimens below are similarly shaped despite of their belonging to different species. Hence the alleged differences in red are merely statistical.
http://luirig.altervista.org/cpm/albums/bot-006/aquilegia-bertolonii11936.jpg
http://4.bp.blogspot.com/_dhmGoQyou-c/S0tzCpbIu_I/AAAAAAAABQM/GM91LIyr77Y/s400/Aquilegia+pyrenaica+blog.jpg
Genuine plants are at pages
http://luirig.altervista.org/schedeit/ae/aquilegia_bertolonii.htm
http://www.fotonatura.org/galerias/flora/198516/
Furthermore, the cultivated plants may be hybrids because it is not in the forces of gardeners to distinguish between the two similar species.
Title: Re: Aquilegia saximontana
Post by: Great Moravian on November 08, 2010, 11:22:00 AM
The information in Flore de France is as follows. It is written in French abbreviations. A funny reading.
It is obviously useless for our purpose. The information in Flora Europaea is useless too.
A.bertoloniiA. pyrenaica subsp. pyrenaica
éperon gént recourbé en crochet `a son extrémité, plus rart simplement arqué ---> -+ droitéperon droit ou un peu arqué, exceptionnellement -+ recourbé en crochet `a son extrémité
t. soit glabres soit -+ poilues-glanduleuses ds leur part. sup.t. glabres ou faiblt poilues-glanduleuses ds leur part. sup.
10-50 cm10-30 cm
French authors include Aquilegia reuteri in Aquilegia bertolonii.
Several Italian authors restrict the latter to Alpi Apuane and regard the former as a member of the Aquilegia einseleana group.
Leaves
Aquilegia bertolonii, Alpi Apuane
lobulis linearibus subacutatis, divergentibus
http://www.actaplantarum.org/floraitaliae/download/file.php?id=3825
Aquilegia pyrenaica
lobulis oblongo-obovatis, truncato-rotundatis, retusis,
subdistantibus

http://www.florasilvestre.es/mediterranea/Ranunculaceae/Aquilegia_pyrenaica.jpg
http://upload.wikimedia.org/wikipedia/commons/6/6b/Aquilegia_pyrenaica.jpg
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on November 09, 2010, 02:01:15 AM
Thanks Moravian for shedding additional light on A. bertolonii and pyrenaica, the color coordinated highlighting of the latin descriptions certainly helps, as does the sequential photo links.

I don't have the Bob Nold book on Aquilegia, anyone have any thoughts they can share about the book.

I keep going back to the Gentes Herbarum 1946 monograph on Aquilegia, even though obviously outdated at this point, it's a truly fine publication packed with information and superb line drawings.  Here are scans from the publication on both A. bertolonii and pyrenaica.
  For sake of keyword searching, there is also a nice line drawing here of A. einseleiana and a flower of A. bernardii.
Title: Re: Aquilegia saximontana
Post by: Great Moravian on November 09, 2010, 11:45:35 AM
I am Great Moravian and not Marovian, otherwise I thank TheOnionMan for further information.
So the leaves should be pilose beneath in Aquilegia bertolonii whereas glabrous beneath in Aquilegia pyrenaica.
Flora Iberica claims subglabrous for the latter.
Sepals are broader in the latter in reality.
Botanists obviously cannot reveal a clear distinctive feature separating the two species.
Title: Re: Aquilegia saximontana
Post by: Maggi Young on November 09, 2010, 12:20:27 PM
I am Great Moravian and not Marovian, otherwise I thank TheOnionMan for further information.

Dear me, so easy to get the names confused, isn't it, even with people?  ;)
Title: Re: Aquilegia saximontana
Post by: Great Moravian on November 09, 2010, 01:44:53 PM
Dear me, so easy to get the names confused, isn't it, even with people?  ;)
No problem. I checked the descriptions by Munz posted above by TheOnionMan. No other differences can be found.
---
Lobuli linear, nearly sharpened, divergent, pilose beneath
         Aquilegia bertolonii
Lobuli oblong-obovate, truncate-rounded, retuse, subdistant,
nearly glabrous beneath
         Aquilegia pyrenaica subsp. pyrenaica
---
Now the alleged Aquilegia jonesii x A. saximontana which is commonly cultivated throughout the world.
(http://www.srgc.org.uk/smf/index.php?action=dlattach;topic=6126.0;attach=248784;image)
Its lobuli of leaves are distinctly truncate-rounded, not resembling the leaves of A. bertolonii.
Therefore Aquilegia pyrenaica subsp. pyrenaica. Nevertheless, the flowers are not ideal.
Perhaps a hybrid.
Title: Re: Aquilegia saximontana
Post by: Maggi Young on November 09, 2010, 02:13:00 PM
Dear me, so easy to get the names confused, isn't it, even with people?  ;)
No problem. I checked the descriptions by Munz posted above by TheOnionMan. No other differences can be found.

Great Moravian.... I was having a little joke about the  Onion Man getting your name wrong.... I'm sorry if the British humour is a little difficult to follow.
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on November 09, 2010, 03:28:42 PM
I am Great Moravian and not Marovian, otherwise I thank TheOnionMan for further information.
So the leaves should be pilose beneath in Aquilegia bertolonii whereas glabrous beneath in Aquilegia pyrenaica.
Sepals are broader in the latter in reality. Botanists obviously cannot reveal a clear distinctive feature separating the two species.

I tend towards dyslexia and frequently transpose letters or numbers, it was just one such slip, sorry about that.  Also note TheOnionMan uses a signature block revealing my identity as Mark McDonough... people on the forum call me McMark to lessen the confusion with omnipresent forumist Mark Smyth.  You can call me Mark or McMark :)

Regarding "Botanists obviously cannot reveal a clear distinctive feature separating the two species", I find this often to be the case, not only in Aquilegia, but in other floras and genera.  Sometimes one has to also know the provenance of wild collected plants or seed to get a clue too, although species descriptions should be able to stand on their own and not rely on provenance for separation.
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on November 10, 2010, 03:43:38 PM
Just some links to selected Calphotos images of Aquilegia scopulorum.

Aquilegia scopulorum - foliage
http://calphotos.berkeley.edu/cgi/img_query?enlarge=0000+0000+1209+2811
http://calphotos.berkeley.edu/cgi/img_query?enlarge=0000+0000+1209+3166
http://calphotos.berkeley.edu/cgi/img_query?enlarge=0000+0000+1209+3169
http://calphotos.berkeley.edu/cgi/img_query?enlarge=0000+0000+1209+3177

Aquilegia scopulorum - flowering plants in the wild, Utah
http://calphotos.berkeley.edu/cgi/img_query?enlarge=0000+0000+0110+1042
http://calphotos.berkeley.edu/cgi/img_query?enlarge=0000+0000+1209+3162
http://calphotos.berkeley.edu/cgi/img_query?enlarge=0000+0000+1209+3168
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on November 11, 2010, 04:49:41 AM
There is continuing presence of the near mythical Aquilegia jonesii x saximontana hybrid, I chanced upon the following article which sheds some light on this entity.

There's an article about this hybrid cross published in the Bulletin of the American Rock Garden Society, Vol. 38, Summer 1980, N0.3, pp. 113-114, "A Dwarf Aquilegia Hybrid", by Trevor Cole, Ottawa Research Station, Agriculture Canada, Ottawa, Ontario.  In the article, Trevor reports on his experience growing seed of A. jonesii which had been collected on the Big Horn Mountains.  In the ensuring years growing and flowering A. jonesii, the only other columbine close by was A. saximontana, reported as 6' (2 m) away.  From seed harvested off his own A. jonesii plants and sown in January 1976, about 40 seedlings were grown on.  In the batch of seedlings, two different forms appeared. Most plants were typical in growth to saximontana, but 5 plants were characteristic of jonesii.  Almost all plants "bloomed profusely in the summer of 1977", and "seed was collected in quite large amounts and sent to the ARGS seed exchange listed as A. jonesii x saximontana".  So indeed, the cross did exist at one point in time.

Surely 33 years hence, with the well known promiscuity of the genus Aquilegia, unless any of these hybrids were grown absolutely isolated, there is almost no chance that anything close to the unadulterated cross between the two species exists today.  It is also important to note, that the seed that was initially distributed, was itself variable, from hybrid plants that already showed a couple different growth characteristics.

Almost everything bearing the name of this cross (and sometimes you can see it listed the other way around) look to be dwarf forms and/or hybrids of A. pyrenaica.
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on November 11, 2010, 05:11:44 AM
This little nugget from the late Linc Foster, with an essential line drawing from Timmy Foster exemplifying the essence of Aquilegia saximontana, a dwarf open spray of leaves and flowers with those spreading or divergent spurs.  This appeared in the Bulletin of the American Rock Garden Society, Vol. 38, Summer 1980, No.3, entitled "Aquilegia saximontana in Connecticut".
Title: Re: Aquilegia saximontana
Post by: maggiepie on November 11, 2010, 03:14:46 PM
Here's another aquilegia I have labelled as saximontana, also grown from seed.

Yes or no?  ??? ??? ???

Sorry the pics aren't very good.
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on November 11, 2010, 08:30:16 PM
Here's another aquilegia I have labelled as saximontana, also grown from seed.

Yes or no?  ??? ??? ???

Sorry the pics aren't very good.

Ding, ding, ding!  Congratulations Helen, give this woman a teddy bear! :o ;D :o   Yes, that looks like A. saximontana, notice the outwardly splayed (divergent) spurs... yours is the first columbine submitted on this topic that looks correct.  Well done!

Where did you get the seed?  Now, there is also a strong possibility this is a hybrid with saximontana, particularly if it came from garden grown seed, but it at least has the right look and is definitely not a flabellata nor a pyrenaica type.
Title: Re: Aquilegia saximontana
Post by: maggiepie on November 11, 2010, 08:38:07 PM
]

Ding, ding, ding!  Congratulations Helen, give this woman a teddy bear! :o ;D :o   Yes, that looks like A. saximontana, notice the outwardly splayed (divergent) spurs... yours is the first columbine submitted on this topic that looks correct.  Well done!

Where did you get the seed?  Now, there is also a strong possibility this is a hybrid with saximontana, particularly if it came from garden grown seed, but it at least has the right look and is definitely not a flabellata nor a pyrenaica type.

Well there's murphy for you, the plant appears to have carked it!!!  :'( :'( :'(

I think the seed came from Alplains, the writing on the label has gone. Grrrrrr ( I did get some from various seed exchanges too)
There are more of them that haven't flowered yet but the plants have stayed tiny and I fear they will not survive winter.



Title: Re: Aquilegia saximontana
Post by: Great Moravian on November 12, 2010, 03:17:40 PM
(http://cs4145.vkontakte.ru/u6450879/93407084/x_86b9ccb8.jpg)(http://www.srgc.org.uk/smf/index.php?action=dlattach;topic=6126.0;attach=253235)
Aquilegia pyrenaica subsp. discolorAquilegia saximontana
Interesting comparison of the two often misidentified plants.
---
Aquilegia pyrenaica subsp. discolor exists in a concolorous variant too.
http://www.asturnatura.com/fotografia/flora/aquilegia-pyrenaica-subsp-discolor/9511.html
Title: Re: Aquilegia saximontana
Post by: Great Moravian on November 12, 2010, 03:29:42 PM
It might be interesting to share information concerning the cultivation of A. saximontana.
It is probably not straightforward assuming common use of easier species instead.
The cultivars of A. flabellata thrive in any non-extreme garden position.
A. pyrenaica subsp. pyrenaica and subsp. discolor require a stony substrate.
Limestone in mountains but anything in the garden.
Title: Re: Aquilegia saximontana
Post by: Great Moravian on November 15, 2010, 11:12:07 AM
Flora iberica
http://www.floraiberica.es/floraiberica/texto/pdfs/01_036_20_Aquilegia.pdf
---
At a sister society-
http://nargs.org/nargswiki/tiki-browse_image.php?imageId=816 A. laramiensis by Lori Skulski
http://nargs.org/smf/index.php?topic=151.0
---
Edited to an effeminate wording by Maggi Young ... sister society.


Title: Re: Aquilegia saximontana
Post by: Lori S. on November 15, 2010, 01:28:24 PM
At a sister society-
http://nargs.org/nargswiki/show_image.php?id=816&scalesize=o
This one, which is my trough plant, is A. laramiensis, not A. saximontana... unless you are saying it is something else?
Title: Re: Aquilegia saximontana
Post by: Great Moravian on November 15, 2010, 02:14:17 PM
At a sister society-
http://nargs.org/nargswiki/show_image.php?id=816&scalesize=o
This one, which is my trough plant, is A. laramiensis, not A. saximontana... unless you are saying it is something else?
Yes, it might be, but A. laramiensis was discussed above too. I would appreciate if you would share the image if possible.

jn
Title: Re: Aquilegia saximontana
Post by: Lori S. on November 16, 2010, 01:59:39 AM
Yes, it might be, but A. laramiensis was discussed above too. I would appreciate if you would share the image if possible.
jn

Yes, it's certainly no problem to share the image.  (I wouldn't post photos on the internet if I felt otherwise!  :))  Since you have linked to the photo, I assume there is no need for me to post it here as well.   I was just surprised to see that the title, etc. from the NARGS photo site did not come across with it.  

Of course, I'm curious now about your comment that "Yes, it might be...".  Are you thinking it is not A. laramiensis, or that it is a similar-looking hybrid of... something?  Isn't A. laramiensis supposed to be very indisposed to hybridizing?  
Well, at any rate, I bought the plant in 1998 and it has been seeding itself around the trough since then.  I don't have any photos from that far back, but the photos from 2004 look the same as the recent ones, anyway...
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on December 01, 2010, 05:12:22 AM
I have the pleasure to post some beautiful photos of Aquilegia saximontana taken by Jane Hendrix, of Breckenridge, Colorado, a purveyor and wonderful photographer of North American native flora, particularly Rocky Mountain alpines.  One can only imagine what it must be like to live and garden at 10,000' elevation, in such close proximity to an amazing alpine flora.  Her extensive photo albums in the following link show many wonderful Rocky Mountain wildflowers and alpines:
http://www.picturetrail.com/hendrix

It is my privilege to show these photos that capture the unique essence and characteristic of the true A. saximontana.  Here again, what we see is the dwarf yet upright billowy foliage of the plant, and the small soft blue and white flared flowers with splayed (divergent) spurs.
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on December 02, 2010, 02:10:34 PM
Following up on the previous post with lovely photos of A. saximontana by Jane Hendrix, Breckenridge, Colorado, USA, Jane has given me permission to post photos of a dwarf form of Aquilegia caerulea found locally among unnamed peaks that surround her 10,000' elevation home.  Rather than paraphrase Jane's description, she describes it well, so here it is:

"Every plant shown was grown from seed I collected at 11,600 feet in the boulder field on the slope of Peak 6 so they are not accidental hybrids.  To be certain this dwarf form [of A. caerulea] was genetically dwarf and not environmentally dwarf, I sited the plants in rich, moist soil.  They all grew lusher, fuller and taller than the ones in the wild but still would be classified as dwarf.  In one photo, you can see the plant's height in relation to the red Darwin tulips (the tulips get to about 20 inches tall in my garden).  The dwarf A. caerulea flowers are as large as the "regular" A. caerulea.  This particular group of seeds produced light blue and white blossoms but since collecting them, I have seen similar dwarf A. caerulea in a nearby rock field with the typical, deeper blue sepals.  When grown side-by-side with the common (pure) A. caerulea, they don't have a chance to hybridize because the dwarf form blooms much earlier than the other one and by the time the tall one has pollen, the little one is making seed."

Thanks Jane for allowing forumists to view this rarely seen variant of Colorado's most famous plant.
Title: Re: Aquilegia saximontana
Post by: maggiepie on December 02, 2010, 02:18:59 PM
A very garden worthy plant, Mark.
Did you get seeds? ;)

Title: Re: Aquilegia saximontana
Post by: mark smyth on December 02, 2010, 02:29:53 PM
and a wink  ;) from me also LOL ;D
Title: Re: Aquilegia saximontana
Post by: TheOnionMan on December 02, 2010, 03:21:35 PM
This dwarf version of A. caerulea might be regarded as A. caerulea var. alpina.

Aquilegia caerulea var. alpina A. Nels. is a recognized variety, in ITIS, USDA, and in Flora of North America (FONA).  The problem is USDA reports var. alpina from Wyoming and Utah (not Colorado) and FONA reports var. alpina only occurring in Wyoming. Using the USDA species distribution, it would be conceivable this variety is also found in adjacent Colorado.
http://plants.usda.gov/java/profile?symbol=AQCOA
http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=233500096

Tip: if searching for information or photos on the Colorado Columbine, use both alternate spellings; caerulea and coerulea.

Aquilegia coerulea photos, it seems that some A. coerulea var. coerulea photos show compact plants:
http://plants.usda.gov/java/largeImage?imageID=aqco_004_ahp.jpg
http://plants.usda.gov/java/largeImage?imageID=aqco_002_ahp.jpg
http://plants.usda.gov/java/largeImage?imageID=aqcac2_001_ahp.tif
Title: Re: Aquilegia saximontana
Post by: Karaba on April 24, 2018, 11:16:12 AM
French authors include Aquilegia reuteri in Aquilegia bertolonii.
Several Italian authors restrict the latter to Alpi Apuane and regard the former as a member of the Aquilegia einseleana group.
It's an old thread but it's the only one dealing with the difference betweeen A. pyrenaica and bertolonii.
I just want to add a few points about the position of french botanists on this duo.

In the last Flora gallica, Aquilegia reuteri is now separated from bertolonii and only A. reuteri is mentionned in the french Alps. In the key, the "only" difference between reuteri and pyrenaica is on the spur, generally hooked (>180°) in reuteri, generally arqued (<90°) in pyrenaica. This caracter might be cheap but if you look at pictures at the following links, you will see that there is a striking difference even if, in the extreme case, spurs can look quite similar (it might also vary with anthesis). The difference on the shape of the spur is also mentionned to differenciate A. alpina and reuteri and this is far less obvious.
The difference in the distribution of the two species do not incite to look for any other big differences and Flora Gallica is mainly a (big) key. In Flore de Coste, the other difference is about the leaves as mentionned by Great Moravian. These differences can be seen in the set of pictures below :
- reuteri :  à folioles profondément incisées-lobées, les supérieures à 3-7 lobes linéaires ou entières = (own poor translation) leaflet deaply incised-lobated, the upper ones with 3-7 linear lobes or entire. In Flore de la France méditerranéenne, there is a precision that the incision is at least 30% of the leaflet.
- pyrenaica : à folioles crénelées ou entières  = leaflet crenulated or entire
There is no difference mentionned in french flora about pilosity of the leaves. Usually mentionned as subglabrous in pyrenaica.

Aquilegia pyrenaica
http://www.photoflora.fr/FiTax.php?NumTaxon=4424 (http://www.photoflora.fr/FiTax.php?NumTaxon=4424)
http://www.florealpes.com/fiche_aquilegiapyrenaica.php (http://www.florealpes.com/fiche_aquilegiapyrenaica.php)
http://www.tela-botanica.org/bdtfx-nn-74964-illustrations (http://www.tela-botanica.org/bdtfx-nn-74964-illustrations)
Aquilegia reuteri
http://www.photoflora.fr/FiTax.php?NumTaxon=4421 (http://www.photoflora.fr/FiTax.php?NumTaxon=4421)
http://www.florealpes.com/fiche_aquilegiabertolonii.php (http://www.florealpes.com/fiche_aquilegiabertolonii.php)
http://www.tela-botanica.org/bdtfx-nn-5725-illustrations (http://www.tela-botanica.org/bdtfx-nn-5725-illustrations)

I looked at pictures of A. bertolonii in Italy (http://luirig.altervista.org/flora/taxa/index1.php?scientific-name=aquilegia+bertolonii (http://luirig.altervista.org/flora/taxa/index1.php?scientific-name=aquilegia+bertolonii)) and, indeed, the caracter on the spur is not flagrant  ;D This is one of the differences that substain the two entity reuteri and bertolonii as it is mentionned here : http://www.floraitaliae.actaplantarum.org/viewtopic.php?t=1065 (http://www.floraitaliae.actaplantarum.org/viewtopic.php?t=1065) and http://www.ambienteinliguria.it/eco3/DTS_PUBBLICAZIONI/20070625/01_Vegetali.pdf (http://www.ambienteinliguria.it/eco3/DTS_PUBBLICAZIONI/20070625/01_Vegetali.pdf) True bertolonii has nearly straight spur while reuteri has hooked spur. I don't made further search about the shape of the leaves between reuteri and bertolonii but it seems that there are some differences (read again http://www.floraitaliae.actaplantarum.org/viewtopic.php?t=1065 (http://www.floraitaliae.actaplantarum.org/viewtopic.php?t=1065)  ;) )
It is also to mention that A. einseleana and bertolonii are together and well differentiated from other european aquilegia in the phylogeny made by Fior (2013) https://www.researchgate.net/publication/235399368_Spatiotemporal_reconstruction_of_the_Aquilegia_rapid_radiation_through_next-generation_sequencing_of_rapidly_evolving_cpDNA_regions (https://www.researchgate.net/publication/235399368_Spatiotemporal_reconstruction_of_the_Aquilegia_rapid_radiation_through_next-generation_sequencing_of_rapidly_evolving_cpDNA_regions) Tjhis result is a bit different from what is described  here  http://www.floraitaliae.actaplantarum.org/viewtopic.php?t=1065 (http://www.floraitaliae.actaplantarum.org/viewtopic.php?t=1065) where they group einseleana and reuteri.
Title: Re: Aquilegia saximontana
Post by: Hannelore on April 25, 2018, 08:42:36 AM
My Aquilegia saximontana from seeds buyed 2013 from Allplains is flowering now. By the my fingers on the second picture you can see how small (and cute)the flower is.

[attachimg=1]
[attachimg=2]
Title: Re: Aquilegia saximontana
Post by: Hannelore on May 04, 2018, 03:41:56 PM
There's something I do not understand. I read about Aquilegia that the stamens ripe not at the same time as the ovary so that the flower cannot be pollinated by itself. So I didn't expect seeds from my Aquilegia saximontana because I have only one plant. But after the flowers wilted, there is seed growing. There was no other Aquilegia flower open in my Garden when they flowered. Was my first information wrong? Or is it not valid for Aquilegia saximontana?
[attach=1]
If someone wishes to try, I'll send him or her seeds as soon as they fall out.

I also thought about the flowering time - it is said that A. saximontana flowers in July-August. But this is in the mountains high. Spring begins there much later, even in the Alps snow was falling still last week. So flowering season here in the middle of Germany can well be much earlier.

What do you think about that?

Best wishes
Hannelore
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